{"id":2273,"date":"2018-12-05T15:24:12","date_gmt":"2018-12-05T06:24:12","guid":{"rendered":"http:\/\/163.180.4.222\/lab\/?p=2273"},"modified":"2018-12-05T15:24:12","modified_gmt":"2018-12-05T06:24:12","slug":"%ec%b1%85-%ec%86%8c%ea%b0%9c-%ec%9d%b4%ec%96%b8-%ec%8a%a4%ed%8a%9c%ec%96%b4%ed%8a%b8%eb%8a%94-%ec%96%b4%eb%96%bb%ea%b2%8c-%ec%bc%80%ec%9d%b4%ed%81%ac%eb%a5%bc-%ec%9e%90%eb%a5%bc%ea%b9%8c","status":"publish","type":"post","link":"https:\/\/biochemistry.khu.ac.kr\/lab\/?p=2273","title":{"rendered":"\ucc45 \uc18c\uac1c – \uc774\uc5b8 \uc2a4\ud29c\uc5b4\ud2b8\ub294 \uc5b4\ub5bb\uac8c \ucf00\uc774\ud06c\ub97c \uc790\ub97c\uae4c?: \uc804\ud654\uae30 \uc904\uc740 \uc65c \ud56d\uc0c1 \uc5c9\ucf1c \uc788\uc744\uae4c?"},"content":{"rendered":"

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\uc804\ud654\uae30 \uc904\uc740 \uc65c \ud56d\uc0c1 \uc5c9\ud0ac\uae4c?\u00a0\ub0b4\uac00 \ub9d0\ud558\ub294 \uc904\uc740 \uae34 \ub098\uc120 \ucf54\uc77c\ucc98\ub7fc \uc0dd\uaca8\uc11c \uc7a1\uc544\ub2f9\uaca8 \uae38\uac8c \ub298\uc77c \uc218 \uc788\ub294 \uc904\uc774\ub2e4. \ubcbd\uc5d0 \uac78\uae30\ub3c4 \ud558\ub294 \uad6c\uc2dd \uc804\ud654\uae30\uc758 \uc1a1\uc218\ud654\uae30\ub294 \uadf8\ub7f0 \uc904\ub85c \ubcf8\uccb4\uc5d0 \uc5f0\uacb0\ub418\uc5b4 \uc788\uc5c8\ub2e4. \ucc98\uc74c \uc804\ud654\uae30\ub97c \uc0ac\uc11c \ubcbd\uc5d0 \uac78\uba74 \uc804\ud654\uae30 \uc904\uc740 \uc608\uc058\uace0 \uc58c\uc804\ud558\uac8c \uc544\ub798\ub85c \ub4dc\ub9ac\uc6b4\ub2e4. \uadf8\ub7ec\ub098 \uba87 \uc8fc\uac00 \uc9c0\ub098\uba74 \uc5c9\ud0a8 \uc790\ub9ac\ub4e4\uc774 \uc0dd\uae34\ub2e4.<\/p>\n

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\uace0\ubb34 \ubc34\ub4dc(\ud2b9\ud788 \uc587\uc740 \ub760\ucc98\ub7fc \uc0dd\uae34 \uace0\ubb34 \ubc34\ub4dc)\uc5d0\uc11c\ub3c4 \ub611\uac19\uc740 \ud604\uc0c1\uc744 \ubcfc \uc218 \uc788\ub2e4. \ub2f9\uc2e0\uc774 \uc591 \ub05d\uc744 \uc5c4\uc9c0\uc640 \uac80\uc9c0\ub85c \uc950\uace0 \uc190\uac00\ub77d\ub4e4\uc744 \ube44\ube44\ub4ef\uc774 \uc6c0\uc9c1\uc774\uba74, \uace0\ubb34 \ubc34\ub4dc\ub294 \uaf2c\uc774\uace0 \uc5c9\ud0a8\ub2e4(<\uadf8\ub9bc 1>). \ub610\ub294 \uc801\ub2f9\ud55c \uae38\uc774\uc758 \uace0\ubb34\uc904 \uc591\ub05d\uc744 \uc7a1\uace0 \uc190\uac00\ub77d\ub4e4\uc744 \ube44\ubcbc\ub3c4 \uac19\uc740 \ud604\uc0c1\uc774 \ubc1c\uc0dd\ud55c\ub2e4.<\/p>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n

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\uace0\ubb34 \ubc34\ub4dc\uc758 \uc288\ud37c\ucf54\uc77c\ub9c1.<\/span><\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n
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\uadf8\ub807\uac8c \uace0\ubb34 \ubc34\ub4dc\ub098 \uc904\uc774 \uaf2c\uc5ec \uc5c9\ud0a4\ub294 \ud589\ub3d9\uc744 \u2018\uc288\ud37c\ucf54\uc77c\ub9c1supercoiling\u2019\uc774\ub77c\uace0 \ud55c\ub2e4. \uc288\ud37c\ucf54\uc77c\ub9c1\uc740 \ud574\uc800 \ud1b5\uc2e0\ucf00\uc774\ube14\uc5d0\uc11c DNA\uae4c\uc9c0 \uacfc\ud559\uc758 \ub2e4\uc591\ud55c \ubd84\uc57c\uc5d0\uc11c \ubc1c\uc0dd\ud55c\ub2e4.<\/p>\n

\ub098\ub294 \uc6b0\ub9ac \uc9d1\uc758 \uc804\ud654\uae30 \uc904\uc774 \uc65c \ub108\ubb34 \uaf2c\uc774\ub294\uc9c0 \uc548\ub2e4. \uc790\uc138\ud55c \ub0b4\uc6a9\uc740 \uc5ec\ub7ec\ubd84\uc758 \uc804\ud654\uae30 \uc904\uacfc \ub2e4\ub97c\uc9c0 \ubab0\ub77c\ub3c4 \uc77c\ubc18\uc801\uc778 \uba54\ucee4\ub2c8\uc998\uc740 \uc544\ub9c8 \ub2e4\ub974\uc9c0 \uc54a\uc744 \uac83\uc774\ub2e4. \uadf8 \uba54\ucee4\ub2c8\uc998\uc740 \uace0\ubb34 \ubc34\ub4dc\ub098 \ucf54\uc77c\uc774 \uc5c9\ud0a4\uac8c \ub9cc\ub4dc\ub294 \uc804\ud615\uc801\uc778 \uba54\ucee4\ub2c8\uc998\uacfc \ub3d9\uc77c\ud558\ub2e4.<\/p>\n

\uc804\ud654\ubca8\uc774 \uc6b8\ub9ac\uba74 \ub098\ub294 \uc624\ub978\uc190\uc73c\ub85c \uc1a1\uc218\ud654\uae30\ub97c \ub4e4\uba74\uc11c \uc804\ud654\uae30 \uc904\uc744 \uaf30\ub2e4. \uadf8\ub2e4\uc74c\uc5d0 \ud1b5\ud654\ud558\uae30 \uc704\ud574 \uc1a1\uc218\ud654\uae30\ub97c \uc67c\uc190\uc73c\ub85c \uc62e\uae30\uba74\uc11c \uc804\ud654\uae30 \uc904\uc744 \ub354 \uaf30\ub2e4. \ud1b5\ud654\uac00 \ub05d\ub098\uba74 \uc67c\uc190\uc73c\ub85c \uc1a1\uc218\ud654\uae30\ub97c \ub0b4\ub824\ub193\uc73c\uba74\uc11c \uc804\ud654\uae30 \uc904\uc744 \ub9c8\uc9c0\ub9c9\uc73c\ub85c \ub354 \uaf30\ub2e4. \uadf8\ub807\uac8c \uc804\ud654\uae30\ub97c \uc4f8 \ub54c\ub9c8\ub2e4 \ub098\ub294 \uc804\ud654\uae30 \uc904\uc744 \ucd1d 360\u00ba\u00a0\uaf30\ub2e4. \ub354\uad6c\ub098 \ub9e4\ubc88 \ub611\uac19\uc740 \ubc29\ud5a5\uc73c\ub85c \uaf30\ub2e4.<\/p>\n

\ub0b4\uac00 \uc1a1\uc218\ud654\uae30\ub97c \uc624\ub978\uc190\uc5d0\ub9cc \ub4e4\uace0 \uc788\ub2e4\uac00 \ub0b4\ub824\ub193\uc73c\uba74 \uc904\uc740 \uaf2c\uc774\uc9c0 \uc54a\uaca0\uc9c0\ub9cc, \uc190\uc744 \ubc14\uafb8\uae30 \ub54c\ubb38\uc5d0 \uc804\ud654\uae30 \uc904\uc740 \uc6b4\uba85\uc801\uc73c\ub85c \uaf2c\uc774\uace0 \ub9cc\ub2e4. \ub9c8\ub2f9\uc5d0\uc11c \uc804\uae30\ub97c \uc4f8 \ub54c \uc774\uc6a9\ud558\ub294 \uc804\uc120\ub3c4 \uac19\uc740 \uc6b4\uba85\uc5d0 \ucc98\ud55c\ub2e4. \ub098\ub294 \uc804\uc120\uc744 \uc4f4 \ub2e4\uc74c\uc5d0 \ub9c8\uce58 \uc0b0\uc545\uc778\ub4e4\uc774 \uc790\uc77c\uc744 \uac10\uc744 \ub54c\ucc98\ub7fc \uac10\ub294\ub370, \uc2dc\uac04\uc774 \uc9c0\ub098\uba74 \uc804\uc120\uc774 \uc810\uc810 \ub354 \uaf2c\uc778\ub2e4. \uc65c \uc774\ub807\uac8c \uaf2c\uc774\ub294\u00a0\uac83\uc77c\uae4c?<\/p>\n

\uc774\ub7f0 \uc885\ub958\uc758 \uc9c8\ubb38\uc744 \ub2e4\ub8e8\ub294 \uc218\ud559 \ubd84\uc57c\ub294 \uc704\uc0c1\uc218\ud559\uc774\ub2e4. \uc704\uc0c1\uc218\ud559\uc740 \u2018\uace0\ubb34\ud310 \uae30\ud558\ud559\u2019, \uc989 \uc5f0\uc18d \ubcc0\ud658\ub4e4\uc758 \uae30\ud558\ud559\uc774\ub77c\uace0 \ud560 \uc218 \uc788\ub2e4. \uc704\uc0c1\uc218\ud559\uc790\ub4e4\uc740 \ub0a9\uc791\ud55c \ub760\ub85c \uace0\ub9ac\ub97c \ub9cc\ub4dc\ub294 \ubc29\ubc95\uc744 \ube44\ud2c0\uae30(\ube44\ud2c0\ub9bc)twist\uc640 \uac10\uae30(\uac10\uae40)writhe, \ub450 \uac00\uc9c0\ub85c \uad6c\ubd84\ud55c\ub2e4.<\/p>\n

\uc774 \ub450 \ubc29\ubc95\uc758 \ucc28\uc774\uc640 \uad00\uacc4\ub97c \uc774\ud574\ud558\ub824\uba74 \ud2bc\ud2bc\ud55c \uc885\uc774\ub85c \uae34 \ub760\ub97c \ub9cc\ub4e4 \ud544\uc694\uac00 \uc788\ub2e4. \uae38\uc774 20cm, \ud3ed 1cm \uc815\ub3c4\uac00 \uc88b\ub2e4. \ud55c \uba74\uc774 \ub2e4\ub978 \uba74\uacfc \uad6c\ubcc4\ub418\ub294 \uac83\uc774 \uc88b\uc73c\ubbc0\ub85c, \ub760\uc758 \ud55c \uba74\uc5d0 \ube68\uac04\uc0c9\uc744 \uce60\ud558\uace0 \ub2e4\ub978 \uba74\uc5d0 \ud30c\ub780\uc0c9\uc744 \uce60\ud55c\ub2e4. \uc544\ub2c8\uba74 \uc560\ucd08\ubd80\ud130 \uc591\uba74\uc758 \uc0c9\uc774 \ub2e4\ub978 \uc885\uc774\ub97c \uc774\uc6a9\ud55c\ub2e4.<\/p>\n

\ub048\uc758 \ud55c \ub05d\uc744 \uc67c\uc190 \uc5c4\uc9c0\uc640 \uac80\uc9c0\ub85c \uc7a1\uc544 \uc67c\uc190\uc740 \uac00\uc2b4\uc5d0 \ubd99\uc774\uace0, \ub048\uc758 \ub2e4\ub978 \ub05d\uc744 \uc624\ub978\uc190 \uc5c4\uc9c0\uc640 \uac80\uc9c0\ub85c \uc7a1\uc544 \uc624\ub978\uc190\uc740 \uc55e\uc73c\ub85c \ubed7\ub294\ub2e4. \uadf8\ub807\uac8c \ub048\uc774 \ub2f9\uc2e0\uc5d0\uac8c\uc11c \uba40\uc5b4\uc9c0\ub294 \ubc29\ud5a5\uc73c\ub85c \ud3c9\ud3c9\ud558\uac8c \ub193\uc774\uac8c \ud55c\ub2e4. \uc7a1\uc744 \ub54c\ub294 \uc5c4\uc9c0\uac00 \uc704\ub85c \uac00\uace0 \ub098\uba38\uc9c0 \uc190\uac00\ub77d\uc740 \uc544\ub798\ub85c \uac00\ub3c4\ub85d \uc7a1\ub294\ub2e4.<\/p>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n

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(a) \uc190\uac00\ub77d \ub458\ub808\ub85c \ub048\uc744 \uac10\ub294\ub2e4. (b) \uc190\uac00\ub77d\uc744 \ube80\ub2e4. (c) \ub048\uc744 \uc7a1\uc544\ub2f9\uaca8 \uac10\uae30(\uac10\uae40)\ub97c \ube44\ud2c0\uae30(\ube44\ud2c0\ub9bc)\ub85c \ubcc0\ud658\ud55c\ub2e4.<\/span><\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n
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\uc774\uc81c \uc624\ub978\uc190\uc744 \uc6c0\uc9c1\uc5ec \ub048\uc744 \uc67c\uc190 \uc911\uc9c0 \ub458\ub808\ub85c \ud55c \ubc14\ud034 \uac10\ub294\ub2e4(<\uadf8\ub9bc 2a>). \uc774\ub54c \uc624\ub978\uc190\uc744 \ud55c \ubc88 \ubc14\uafd4 \uc7a1\uc544\uc57c \ud558\ub294\ub370, \ubcf5\uc7a1\ud558\uac8c \ub4e4\ub9b4\uc9c0 \ubaa8\ub974\uc9c0\ub9cc \uc9c1\uc811 \ud574\ubcf4\uba74 \uac04\ub2e8\ud558\ub2e4. \uadf8\ub807\uac8c \uac10\uc740 \ub2e4\uc74c\uc5d0\ub294 \uc67c\uc190 \uc911\uc9c0\ub97c \ube7c \ube48 \uace0\ub9ac\ub9cc \ub0a8\uac8c \ud55c\ub2e4(<\uadf8\ub9bc 2b>).<\/p>\n

\uc774\ub85c\uc368 \ub2f9\uc2e0\uc740 \uc6d0\ub798 \ud3c9\ud3c9\ud588\ub358 \ub048\uc5d0 1\ud68c\uc758 \uac10\uae40\uc744 \uc9d1\uc5b4\ub123\uc5c8\ub2e4. \ub9cc\uc77c \uc885\uc774\uac00 \ubb34\ud55c\ud788 \ud0c4\ub825\uc801\uc774\ub77c\uba74(\uc2e4\uc81c \uc885\uc774\ub294 \uadf8\ub807\uc9c0 \uc54a\uc9c0\ub9cc \uc704\uc0c1\uc218\ud559\uc790\ub4e4\uc758 \uc885\uc774\ub294 \uadf8\ub807\ub2e4) \uac10\uae34 \ubd80\uc704\ub97c \ub0a9\uc791\ud558\uac8c \ub9cc\ub4e4\uc5b4 \ud3c9\uba74\uc5d0 \ub193\uc744 \uc218 \uc788\uc744 \uac83\uc774\ub2e4. \ubb3c\ub860 \uadf8\ub7ec\ub824\uba74 \ub048\uc774 \uc790\uae30 \uc790\uc2e0\uacfc \uacb9\uccd0\uc9c0\uaca0\uc9c0\ub9cc\u00a0\ub9d0\uc774\ub2e4.<\/p>\n

\uadf8\ub7f0\ub370 <\uadf8\ub9bc 2b>\uc758 \uc0c1\ud669\uc73c\ub85c \ub3cc\uc544\uac00 \ub450 \uc190\uc744 \ucc9c\ucc9c\ud788 \ubc8c\ub824\ubcf4\uc790. \ub048\uc740 <\uadf8\ub9bc 2c>\ucc98\ub7fc \ubcc0\ud615\ub41c\ub2e4. \uc774\uac83\uc740 \uac10\uae40\uc774 \uc544\ub2c8\ub77c \ube44\ud2c0\ub9bc\uc774\ub2e4. \ub2f9\uc2e0\uc740 \ub048\uc744 \ub2f9\uc2e0 \uc55e\uc5d0 \uac00\ub85c\ub85c \ub193\uace0 \uc67c\ucabd \ub05d\uc744 \uadf8\ub300\ub85c \uc720\uc9c0\ud558\uba74\uc11c \uc624\ub978\ucabd \ub05d\uc744 360\u00ba \ube44\ud2c0\uc5b4\uc11c \ub3d9\uc77c\ud55c \uacb0\uacfc\ub97c \uc5bb\uc744 \uc218 \uc788\ub2e4. \uadf8\ub7ec\ubbc0\ub85c \uc704\uc0c1\uc218\ud559\uc801\uc73c\ub85c \ubcfc \ub54c \uac10\uae30(\uac10\uae40)\ub294 \ube44\ud2c0\uae30(\ube44\ud2c0\ub9bc)\ub85c \ubcc0\ud615\ub420 \uc218 \uc788\ub2e4\ub294 \uac83\uc744 \uc54c \uc218 \uc788\ub2e4.<\/p>\n

\uc774\ub54c \uc911\uc694\ud55c \uc804\ubb38\uc801\uc778 \uc0ac\ud56d\uc774 \ud558\ub098 \uc788\ub2e4. \uac10\uae40\uacfc \ube44\ud2c0\ub9bc\uc740 \ub458 \ub2e4 \ubc29\ud5a5\uc774 \uc788\ub2e4. \ub458 \ub2e4 \u2018\uc591positive\u2019\uc774\uac70\ub098 \u2018\uc74cnegative\u2019\uc77c \uc218 \uc788\ub2e4. \ubb34\uc5c7\uc774 \uc591\uc758 \uac10\uae40(\ube44\ud2c0\ub9bc)\uc774\uace0 \ubb34\uc5c7\uc774 \uc74c\uc758 \uac10\uae40(\ube44\ud2c0\ub9bc)\uc778\uc9c0 \ud310\uc815\ud558\ub294 \uac83\uc740 \uadf8\ub2e4\uc9c0 \uc5b4\ub835\uc9c0 \uc54a\uc740 \uc77c\uc774\ub098, \ub098\ub294 \uc138\ubd80\uc0ac\ud56d\uc73c\ub85c \ub3c5\uc790\ub4e4\uc744 \ubd80\ub2f4\uc2a4\ub7fd\uac8c \ud558\uace0 \uc2f6\uc9c0\ub294 \uc54a\ub2e4. \uadf8\ub7ec\ub2c8 \uc5ec\ub7ec\ubd84\uc740 \uacf0\ub3cc\uc774 \ud478\uac00 \uc67c\ucabd\uacfc \uc624\ub978\ucabd\uc744 \ub9de\ud788\ub294 \ubb38\uc81c\ub97c \ud574\uacb0\ud560 \ub54c \uc4f4 \ubc29\ubc95\uc744 \uc368\uc57c \ud55c\ub2e4. \ub140\uc11d\uc740 \uc77c\ub2e8 \uc5b4\ub290 \ubc1c\uc774 \uc624\ub978\ubc1c\uc778\uc9c0 \uc54c\uba74, \ub2e4\ub978 \ubc1c\uc740 \uc67c\ubc1c\uc774\ub77c\ub294 \uc810\uc744 \uae68\ub2ec\uc558\ub2e4. \ub2e4\ub9cc \ucc98\uc74c\uc5d0 \uc5b4\ub290 \ubc1c\uc774 \uc624\ub978\ubc1c\uc778\uc9c0 \ubaa8\ub978\ub2e4\ub294 \uc810\uc774 \ubb38\uc81c\uc600\ub2e4.<\/p>\n

\uc774\ub97c \uc6b0\ub9ac\uc758 \ub17c\uc758\uc5d0 \uc801\uc6a9\ud558\uba74 \uc774\ub807\ub2e4. \ub2f9\uc2e0\uc774 \ud2b9\uc815\ud55c \uac10\uae40\uc774\ub098 \ube44\ud2c0\ub9bc\uc744 \uc591\uc73c\ub85c \uacb0\uc815\ud558\uba74, \uadf8\uac83\uc758 \uac70\uc6b8\uc0c1\uc740 \uc74c\uc774\ub2e4. \uc27d\uac8c \uc2dc\uc791\ud558\ub294 \ubc29\ubc95\uc740 <\uadf8\ub9bc 2b>\uc758 \uac10\uae40\uc744 \uc591\uc758 \uac10\uae40\uc73c\ub85c, <\uadf8\ub9bc 2c>\uc758 \ube44\ud2c0\ub9bc\uc744 \uc74c\uc758 \ube44\ud2c0\ub9bc\uc73c\ub85c \uc120\uc5b8\ud558\ub294 \uac83\uc774\ub2e4. \uadf8\ub807\ub2e4\uba74 \ube44\ud2c0\ub9bc \uc218\ub294 \uac10\uae40 \uc218\uc5d0 \uc74c\uc758 \ubd80\ud638\ub97c \ubd99\uc778 \uac83\uacfc \uac19\ub2e4. \uc774\ub807\uac8c \ud558\uba74 \uac04\ub2e8\ud55c \ubc29\uc815\uc2dd T+W=0\uc744 \uc5bb\uc744 \uc218 \uc788\ub2e4. \uc774\ub54c T\ub294 \ube44\ud2c0\ub9bc \uc218, W\ub294 \uac10\uae40 \uc218\ub2e4. \ubd80\ud638\ub97c \ub2e4\ub974\uac8c \uc815\ud55c\ub2e4\uba74 T-W=0\uc744 \uc5bb\uc744 \uc218 \uc788\uc744 \uac83\uc774\ub2e4. \uc5b4\ub290 \ucabd\uc774\ub4e0 \ud0c0\ub2f9\ud558\uc9c0\ub9cc, \ubc18\ub4dc\uc2dc \uc77c\uad00\ub418\uac8c \ud55c\ucabd\uc744 \uc120\ud0dd\ud574\uc57c \ud55c\ub2e4.<\/p>\n

\ub2e4\uc2dc \ucc98\uc74c\uc758 \ud3c9\ud3c9\ud55c \ub048\uc73c\ub85c \ub3cc\uc544\uac00\uc11c, \uc774\ubc88\uc5d0\ub294 \ub048\uc744 \uc67c\uc190 \uc911\uc9c0 \ub458\ub808\ub85c \ub450 \ubc14\ud034 \uac10\ub294\ub2e4. \ub048\uc740 (+)2\ud68c\uc758 \uac10\uae40\uc744 \uc5bb\uc5c8\ub2e4. \uadf8 \uc0c1\ud0dc\uc5d0\uc11c \uc591\uc190\uc744 \ubc8c\ub9ac\uba74, 2\ud68c \uac10\uae40\uc740 2\uc911 \ube44\ud2c0\ub9bc(720\u00ba \ube44\ud2c0\ub9bc)\uc73c\ub85c \ubc14\ub010\ub2e4. \uc774\ucc98\ub7fc (+)2 \uac10\uae40\uc740 (-)2 \ube44\ud2c0\ub9bc\uc774 \ub420 \uc218 \uc788\ub2e4. \uacbd\uc6b0\uc5d0 \ub530\ub77c\uc11c\ub294 (+)2 \uac10\uae40\uc774 (+)1 \uac10\uae40 \ub354\ud558\uae30 (-)1 \ube44\ud2c0\ub9bc\uc774 \ub420 \uc218\ub3c4 \uc788\ub2e4. 3\ud68c \ub610\ub294 4\ud68c \uac10\uae40\uc73c\ub85c\ub3c4 \uc2e4\ud5d8\ud574\ubcf4\ub77c. \ud2b9\uc815 \ud69f\uc218\uc758 (\uc591\uc758) \uac10\uae40\uc740 \ub3d9\uc77c\ud55c \ud69f\uc218\uc758 (\uc74c\uc758) \ube44\ud2c0\ub9bc\uc73c\ub85c \ubcc0\ud658\ub420 \uc218 \uc788\uc74c\uc744 \uc54c\uac8c \ub41c\ub2e4.<\/p>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n

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(a) \ud558\ub098\uc758 \uac10\uae40\uc774 \ud558\ub098\uc758 \ube44\ud2c0\ub9bc\uc73c\ub85c \ub41c\ub2e4. (b) \uac19\uc740 \uacfc\uc815\uc744 \ubc18\ubcf5\ud558\uba74, \uc784\uc758\uc758 \uac1c\uc218\uc758 \uac10\uae40\ub4e4\uc744 \ub3d9\uc77c\ud55c \uac1c\uc218\uc758 \ube44\ud2c0\ub9bc\ub4e4\ub85c \ubcc0\ud658\ud560 \uc218 \uc788\ub2e4.<\/span><\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n
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\uc2e4\uc81c\ub85c \uc6b0\ub9ac\ub294 \uc774\ub7f0 \uc0ac\uc2e4\uc744 \uc99d\uba85\ud560 \uc218 \uc788\ub2e4. <\uadf8\ub9bc 3a>\ub294 \uc5b4\ub5bb\uac8c \uc591\uc758 \uac10\uae40 1\ud68c\uac00 \uc74c\uc758 \ube44\ud2c0\ub9bc 1\ud68c\ub85c \ubcc0\ud658\ub418\ub294\uc9c0 \ubcf4\uc5ec\uc900\ub2e4. \uc774\ub54c \ub048\uc758 \uc591\ub05d\uc740 \uace0\uc815\ub41c \ubc29\ud5a5\uc744 \uc720\uc9c0\ud574\uc57c \ud55c\ub2e4. \ub9c8\uce58 \ub2f9\uc2e0\uc758 \uc190\uac00\ub77d\uc5d0 \ub20c\ub824 \ucc45\uc0c1\uc5d0\uc11c \ub5a8\uc5b4\uc9c8 \uc218 \uc5c6\ub294 \uac83\ucc98\ub7fc \ub9d0\uc774\ub2e4. <\uadf8\ub9bc 2b>\ub294 (3\uac1c\uc758) \uac10\uae40\ub4e4\uc758 \uc5f4\uc744 \ubcf4\uc5ec\uc900\ub2e4. \uc6b0\ub9ac\ub294 \uc774 \uc5f4\uc744 \ub9c8\uc74c\uc18d\uc73c\ub85c 2\uac1c\uc758 \u2018\uacbd\uacc4\ub4e4\u2019\uc744 \ub530\ub77c \ubd84\ud560\ud558\uc5ec \ub2e8\uc77c\ud55c \uac10\uae40 3\uac1c\ub85c \ub098\ub20c \uc218 \uc788\ub2e4.<\/p>\n

\uadf8\ub2e4\uc74c\uc5d0\ub294 \uac01\uac01\uc758 \ub2e8\uc77c\ud55c \uac10\uae40\uc744, \uacbd\uacc4\uc120\ub4e4\uc744 \ucc45\uc0c1\uc5d0 \ubd99\uc778 \ucc44\ub85c, \ube44\ud2c0\ub9bc\uc73c\ub85c \ubcc0\ud658\ud560 \uc218 \uc788\ub2e4. \uc774\ub54c \uacbd\uacc4\ub4e4\uc758 \uc815\ud5a5\uc740 \ubcc0\ud558\uc9c0 \uc54a\uc73c\ubbc0\ub85c, \uadf8 3\uac1c\uc758 \ube44\ud2c0\ub9bc\uc740 \uc11c\ub85c \u2018\ubd99\uc5b4\uc11c\u2019 3\uc911-\ube44\ud2c0\ub9bc\uc774 \ub41c\ub2e4. \ub2f9\uc5f0\ud55c \ub9d0\uc774\uc9c0\ub9cc, \uc774 \uc608\uc5d0\uc11c \uac10\uae40\uc774\ub098 \ube44\ud2c0\ub9bc\uc774 3\uac1c\uc778 \ub370\ub294 \ud2b9\ubcc4\ud55c \uc774\uc720\uac00 \uc5c6\ub2e4. \uadf8\ub7ec\ubbc0\ub85c \uc6b0\ub9ac\ub294 \ud2b9\uc815\ud55c \uac1c\uc218\uc758 \uc591\uc758 \uac10\uae40\uc744 \uac00\uc9c4 \ub048\uc740 \ub3d9\uc77c\ud55c \uac1c\uc218\uc758 \uc74c\uc758 \ube44\ud2c0\ub9bc\uc744 \uac00\uc9c4 \ub048\uc73c\ub85c \ubcc0\ud658\ub420 \uc218 \uc788\ub2e4\ub294 \uacb0\ub860\uc744 \ub0b4\ub9b0\ub2e4. \uc55e\uc5d0\uc11c \uc5b8\uae09\ud588\ub4ef\uc774, T+W=0\uc774\ub2e4.<\/p>\n

\uc5bc\ud54f \uc0dd\uac01\ud558\uba74, \ub0a9\uc791\ud55c \ub048\uc740 \uace0\ubb34 \ubc34\ub4dc\uc640 \ub2ec\ub77c \ubcf4\uc778\ub2e4. \uadf8\ub7ec\ub098 \ub048\uc758 \uc911\uc2ec\uc120\uc744 \ub530\ub77c \uace0\ubb34 \ubc34\ub4dc\uac00 \ub458\ub824 \uc788\ub2e4\uace0 \uc0c1\uc0c1\ud558\uba74 \ub048\uc758 \uc288\ud37c\ucf54\uc77c\ub9c1\uc744 \uc27d\uac8c \uc774\ud574\ud560 \uc218 \uc788\ub2e4. \ub2f9\uc2e0\uc774 \ub048\uc758 \ud55c \ub05d\uc744 \ube44\ud2c0\uba74, \uc0c1\uc0c1\uc758 \ubc34\ub4dc\ub3c4 \ube44\ud2c0\ub9b4 \uac83\uc774\uba70, \ubc34\ub4dc\uc758 \ube44\ud2c0\ub9bc \ud69f\uc218\ub294 \ub2f9\uc2e0\uc774 \ub048\uc744 \ube44\ud2bc \ud69f\uc218\uc640 \ub3d9\uc77c\ud560 \uac83\uc774\ub2e4. \ub9cc\uc77c \ub2f9\uc2e0\uc774 \ub048\uc744 \ud33d\ud33d\ud558\uac8c \uace0\uc815\ud55c\ub2e4\uba74, \ub048\uc740 \uadf8\uc800 \ube44\ud2c0\ub9b4 \ubfd0\uc774\uc9c0\ub9cc, \ub9cc\uc77c \ub048\uc758 \uc591\ub05d\uc774 \uc6c0\uc9c1\uc774\ub3c4\ub85d \ud5c8\uc6a9\ud55c\ub2e4\uba74, \ub048\uc740 \uac10\uae30\ub294 \ud3b8\uc744 \uc120\ud638\ud560 \uac83\uc774\uace0 \uc288\ud37c\ucf54\uc77c\ub9c1\uc774 \ub098\ud0c0\ub0a0 \uac83\uc774\ub2e4.<\/p>\n

\ub048\uc774 \uac10\uae40\uc744 \uc120\ud638\ud558\ub294 \uc774\uc720\ub294 \ub048\uc774 \uc57d\uac04 \ud0c4\uc131\uc801\uc774\ub77c\ub294 \uc0ac\uc2e4\uacfc \uad00\ub828\ub41c\ub2e4. \ub048\uc740 \u2018\uace0\ubb34 \ubc34\ub4dc\u2019\ucc98\ub7fc \ud0c4\uc131\uc801\uc774\uc9c0\ub294 \uc54a\uc9c0\ub9cc, \uc720\uc5f0\ud558\uba70 \ud718\uc5b4\uc9c0\uba74 \ubcf5\uc6d0\ub825\uc744 \ubc1c\ud718\ud55c\ub2e4\ub294 \uc758\ubbf8\uc5d0\uc11c \ud0c4\uc131\uc801\uc774\ub77c\uace0 \ud560 \uc218 \uc788\ub2e4. \ub2f9\uc2e0\uc774 \ub048\uc744 \uad6c\ubd80\ub9ac\uba74 \uad6c\ubd80\ub9b4\uc218\ub85d, \ub048\uc740 \ub354 \uac15\ud558\uac8c \ub2e4\uc2dc \ud3b4\uc9c0\ub824\uace0 \ud55c\ub2e4. \ub048\uc774 \ube44\ud2c0\ub9bc\ubcf4\ub2e4 \uac10\uae40\uc744 \ub354 \uc120\ud638\ud558\ub294 \ud604\uc0c1\uc740 1883\ub144\uc5d0 A. G. \uadf8\ub9b0\ud790A. G. Greenhill\uc5d0 \uc758\ud574 \ucc98\uc74c\uc73c\ub85c \uc124\uba85\ub418\uc5c8\ub2e4. \uadf8\ub294 \uac10\uae34 \ubaa8\uc591\uc774 \uadf8\uc5d0 \ub300\uc751\ud558\ub294 \ube44\ud2c0\ub9b0 \ubaa8\uc591\ubcf4\ub2e4 \ud0c4\uc131\uc5d0\ub108\uc9c0\uac00 \ub354 \uc801\ub2e4\ub294 \uc0ac\uc2e4\uc744 \uc99d\uba85\ud588\ub2e4. \uc774 \uc0ac\uc2e4\uc740 \uc2ec\uc9c0\uc5b4 \uc885\uc774\ub048\uc5d0\ub3c4 \uc801\uc6a9\ub41c\ub2e4.<\/p>\n

\uc2e4\uc81c\ub85c \uc2e4\ud5d8\uc744 \ud574\ubcf4\uba74 \ud655\uc778\ud560 \uc218 \uc788\ub2e4. \uc885\uc774\ub048\uc744 \ud33d\ud33d\ud788 \uc7a1\uc544\ub2f9\uaca8 \uc5d0\ub108\uc9c0\ub97c \ubd80\uc5ec\ud558\uc9c0 \uc54a\ub294\ub2e4\uba74, \uc885\uc774\ub048\uc740 \uac10\uae30\ub294 \ud3b8\uc744 \uc120\ud638\ud55c\ub2e4. \uadf8\ub9b0\ud790\uc740 \ub9cc\uc77c \ubb34\ud55c\ud788 \uae34 \ub9c9\ub300\uae30\uac00 \u2018\ubb34\ud55c\uc810\u2019\uc5d0\uc11c \ubc1c\ud718\ub418\ub294 \ud798\uc5d0 \uc758\ud574 \ube44\ud2c0\ub9ac\uba74 \ub098\uc120helix \ubaa8\uc591\uc73c\ub85c \ud718\uc5b4\uc9c4\ub2e4\ub294 \uac83\ub3c4 \uc99d\uba85\ud588\ub2e4. 1990\ub144\uc5d0 J. \ucf54\uc778J. Coyne\uc740 \uc774 \ub098\uc120\uc774 \uc2e0\uc18d\ud558\uac8c \ub2e8\uc77c\ud55c \ube44\ud2c0\ub9bc\uc73c\ub85c \uad6d\uc18c\ud654\ub418\uba70, \ub9c9\ub300\uac00 \uacb0\uad6d \uc218\ucd95\ud558\uc5ec \uadf8 \ube44\ud2c0\ub9bc\uc740 \uad6d\uc9c0\ud654\ub41c \uc791\uc740 \uace0\ub9ac(\uac10\uae40)\ub85c \ubc14\ub010\ub2e4\ub294 \uac83\uc744 \uc99d\uba85\ud588\ub2e4. \ub9cc\uc77c \ub9c9\ub300\uac00 \ub354 \uc218\ucd95\ud558\ub3c4\ub85d \ud5c8\uc6a9\ud558\uba74, \uadf8 \uace0\ub9ac\ub294 \uc810\uc810 \ub354 \ub9ce\uc740 \uac1c\uc218\uc758 \uac10\uae40\uc744 \uc5bb\ub294\ub2e4.<\/p>\n

\uc624\uc2a4\ud2b8\ub9ac\uc544\uc758 \uc218\ud559\uc790 3\uba85\u2014D. M. \uc2a4\ud140\ud504D. M. Stump, W. B. \ud504\ub808\uc774\uc800W. B. Fraser, K. E. \uac8c\uc774\uce20K. E. Gates\u2014\uc740 \ub354 \ud604\uc2e4\uc801\uc778 \ubaa8\ud615\ud654 \uc804\uc81c\ub4e4\uc744 \uc368\uc11c \ube44\ud2c0\ub9b0 \ub9c9\ub300\uc758 \ud0c4\uc131 \uc774\ub860\uc744 \ubd84\uc11d\ud588\ub2e4. \uadf8\ub4e4\uc740 \uc288\ud37c\ucf54\uc77c\uc758 \uc815\ud655\ud55c \ubaa8\ud615\uc5d0 \uad00\ud55c \uad6c\uccb4\uc801\uc778 \uacf5\uc2dd\ub4e4\uc744 \ubc1c\uacac\ud588\ub294\ub370, \uadf8 \uacf5\uc2dd\ub4e4\uc740 \ud2b9\ud788 \ud574\uc800\ucf00\uc774\ube14\uc744 \ub193\ub294 \uae30\uc220\uc790\ub4e4\uc5d0\uac8c \uc720\uc6a9\ud558\ub2e4. \uc65c\ub0d0\ud558\uba74\u00a0\uadf8\ub7f0 \uc720\ud615\uc758 \ube44\ud2c0\ub9bc\uc740 \ud574\uc800\ucf00\uc774\ube14 \uacf5\uc0ac\uc5d0\uc11c \ud754\ud788 \ubc1c\uc0dd\ud558\ub294 \ubb38\uc82f\uac70\ub9ac\uc774\uae30 \ub54c\ubb38\uc774\ub2e4.<\/p>\n

\uc804\ud654\uae30 \uc904\uc740 \uc6d0\ub9ac\uc801\uc73c\ub85c \ub354 \ubcf5\uc7a1\ud558\ub2e4. \uc65c\ub0d0\ud558\uba74 \uc804\ud654\uae30 \uc904\uc740 \ucc98\uc74c\ubd80\ud130 \ub098\uc120\uc774\uba70 \uc774\ubbf8 \ube44\ud2c0\ub824 \uc788\uae30(\ub610\ub294 \uad00\uc810\uc744 \ub2ec\ub9ac\ud558\uba74, \uac10\uaca8 \uc788\uae30) \ub54c\ubb38\uc774\ub2e4. \ud558\uc9c0\ub9cc \ub0a9\uc791\ud55c \ub048\uacfc \ub9c8\ucc2c\uac00\uc9c0\ub85c \uc804\ud654\uae30 \uc904 \uc5ed\uc2dc \ube44\ud2c0\ub9bc\uc744 \uac10\uae40\uc73c\ub85c \ubcc0\ud658\ud55c\ub2e4. \uc801\uc5b4\ub3c4 \ub2f9\uc2e0\uc774 \uc804\ud654\uae30 \uc904\uc774 \uc6d0\ub798\ubd80\ud130 \uc9c0\ub2cc \ub098\uc120\ud615 \uaf2c\uc784\uc744 \ud480\uc5b4\ubc84\ub9ac\uc9c0 \uc54a\ub294\ub2e4\uba74(\uadf8 \uaf2c\uc784\uc744 \ud480\uc5b4\ubc84\ub9ac\ub294 \uc0ac\ub78c\uc740 \uac70\uc758 \uc5c6\uc744 \uac83\uc774\ub2e4), \uadf8\ub7f0 \ubcc0\ud658\uc774 \uc77c\uc5b4\ub098\uae30 \ub9c8\ub828\uc774\ub2e4. \uc804\ud654\uae30 \uc904\uc758 \ub098\uc120 \ub0b4\ubd80\uc5d0 \uad75\uc740 \uc904\uc774 \ub07c\uc6cc\uc838 \uc788\uace0, \uadf8 \uad75\uc740 \uc904 \uc18d\uc5d0 \ub0a9\uc791\ud55c \ub048\uc774 \ub4e4\uc5b4 \uc788\ub2e4\uace0 \uc0c1\uc0c1\ud574\ubcf4\ub77c. \uc804\ud654\uae30 \uc904\uc774 \ube44\ud2c0\ub9ac\uba74, \uc904\ub3c4 \ube44\ud2c0\ub9ac\uace0 \ub530\ub77c\uc11c \ub048\ub3c4 \ube44\ud2c0\ub9b4 \uac83\uc774\ub2e4.<\/p>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n<\/div>\n

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\uc6d0\ubb38: \uc5ec\uae30<\/a>\ub97c \ud074\ub9ad\ud558\uc138\uc694~<\/p>\n

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 <\/p>\n","protected":false},"excerpt":{"rendered":"

    \uc804\ud654\uae30 \uc904\uc740 \uc65c \ud56d\uc0c1 \uc5c9\ud0ac\uae4c?\u00a0\ub0b4\uac00 \ub9d0\ud558\ub294 \uc904\uc740 \uae34 \ub098\uc120 \ucf54\uc77c\ucc98\ub7fc \uc0dd\uaca8\uc11c \uc7a1\uc544\ub2f9\uaca8 \uae38\uac8c \ub298\uc77c \uc218 \uc788\ub294 \uc904\uc774\ub2e4. \ubcbd\uc5d0 \uac78\uae30\ub3c4 \ud558\ub294 \uad6c\uc2dd \uc804\ud654\uae30\uc758 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